A “cupule” is a noun that refers to a small cup-like structure or sac that is often found in plants, particularly in the context of botanical morphology. It can also refer to the calyx or protective covering that encases the fruit of certain plants, such as acorns, where the cupule is the cup-like base that holds the nut. In some contexts, it may also refer to a small, hollowed-out area or indentation in various biological or geological formations.